Annual or perennial herbs, undershrubs or shrubs. Leaves simple, entire or lobed,
mostly cordate at base, acute or acuminate at apex, palminerved without nectaries.
Flowers axillary, solitary, sometimes in lax panicles by reduction or decrescence of upper
leaves, rarely in lax corymbose racemes; pedicels jointed in the upper half. Epicalyx
absent. Calyx usually campanulate; lobes 5, divided to the middle or below. Corolla
usually yellow, orange, white or pink, rarely with a dark purple centre, rotate, campanu-
late. Staminal column shorter than petals, much widened at base. Carpels and style-
branches 5 - 40, styles filiform to clavate, capitate stigmatose at apex. Schizocarps
globular, campanulate, rarely discoid; mericarps 5 -
40, dehiscent, follicular, flattened-
reniform, round, acuminate or biaristate at apex, often falling leaving a slender truncate,
columella. Seeds 2 -
9 in each mericarp, reniform to subreniform, upper ones ascending;
lower ones pendulous or horizontal, finally falling out of mericarp.
In tropical and subtropical regions of the World, ca 150 species; 12 in India.
KEY TO THE SPECIES
1a. Carpels 5 -
12
2
b. Carpels more than 15
10
2a. Carpels 5
3
b. Carpels 8 -
12
4
3a. Calyx lobes 7 - 10 mm long connate at base; staminal column ca 5.5 mm long, filaments ca 15 mm long
b. Schizocarp ovoid to subcylindric, more than 1 cm long
7
6a. Pedicels shorter than petioles; mericarps awned; seeds ca 2 mm across, minutely puberulous; leaves glabrescent or minutely stellate-pubescent above and densely stellate-pubescent beneath
10a. Stems, petioles and pedicels with dense long patent simple hairs, few minute stellate hairs and short
viscid glandular hairs; corolla orange yellow with a purple centre
b. Stems, petioles and pedicels densely stellate-pubescent with few simple hairs; corolla with or without
purple centre
11
11a. Corolla yellow to pale yellow without purple centre, calyx lobes ovate to lanceolate; mericarps dorsally
and ventrally stellate-hairy; leaves ovate-cordate
12
b. Corolla yellow with deep brown center; calyx lobes deltoid to ovate; mericarps hairy on dorsal margin;
leaves orbicular to rotund-cordate