Herbs, shrubs or trees, often climbers, usually prickly. Leaves simple, alternate,
lobed or entire; petiolate. Flowers minute, bisexual in much branched axillary or
terminal umbellate cymes. Sepals 5, connate at base. Petals 5 with a narrow claw and
a hooded limb which usually enter the staminal cup and cover the anther lobes,
sometimes with two strap-shaped appendages on both sides of the hood and a long
process. Staminal cup with an inner serries of 5 stamens and an outer series of 5
staminodes alternating with stamens; anther lobes reniform, divergent, extrorse; staminodes ovate, acute. Ovary small, 5-loculed; ovules 2 in each locule; styles entire or 5-fid.
Capsules globose, spiny, or prickly, septicidally 5-valved, valves breaking away from the
central column. Seed 1 in each locule, exalbuminous; cotyledons folded.
Tropical America, Africa, Mascarene Islands, Tropical Asia, West Polynesia, ca 50
species; 4 in India.
KEY TO THE SPECIES
1a. Leaves entire
2
b. Leaves lobed
3
2a. Leaves longer than broad, up to 7 cm long, membranous, dentate