Botanical Survey of India | Flora of India

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1a. Woody climbers, stem with recurved spines in leaf axils 2
b. Trees or erect shrubs, stem with straight spines in leaf axils 3
2a. Mature leaves trifoliolate on long and stiff petioles; inflorescence racemose or paniculate 5. Luvunga
b. Mature leaves unifoliolate (or simple) on short, slender and twisted petioles; inflorescence more 9. Paramignya
3a. Fruits larger, up to 15 cm across, with a thick, hard and woody pericarp; seeds woolly-pubescent 4
b. Fruits small to large (up to 12 cm across or more as in some species of Citrus), with a thin or soft skin or a coriaceous, glandular pericarp; seeds glabrous 5
4a. Leaves imparipinnate; petioles and rachis marginate or narrowly winged; leaflets 5 or 7, smaller (ca 3 x 2 cm); stamens 10 - 12; filaments free, woolly-pubescent below; ovary incompletely septate, unilocular with 5 or 6 parietal placentae 4. Limonia
b. Leaves pinnately trifoliolate; petioles wingless; leaflets up to 13 x 7 cm; stamens many (30 - 40); filaments irregularly united at base, glabrous; ovary 8 - 12 (-20)-locular with many, biseriate ovules 1. Aegle
5a. Fruits thin-skinned soft berries; locules filled with mucilaginous gum without pulp-vesicles 6
b. Fruits with an outer leathery glandular pericarp; locules with or without pulp-vesicles, semidry or juicy 7
6a. Leaves simple; berries purplish, turning black when ripe 11. Severinia
b. Leaves trifoliolate; berries deep reddish when ripe 12. Triphasia
7a. Leaves imparipinnate; leaflets smaller, sessile, opposite; petiole and rachis winged; fruits bluish-black when ripe 7. Naringi
b. Leaves simple, or tri- or unifoliolate; leaflets larger, petiolulate; petioles wingless or narrow to broadly winged; fruits orange or lemon-like when ripe 8
8a. Leaves simple, thick, fleshy with rather indistinct venation; fruits without pulp-vesicles 9
b. Leaves uni- or trifoliolate, thin, moderately coriaceous, with rather prominent venation; petioles articulate; fruits with rudimentary or well developed pulp-vesicles 10
9a. Inflorescence of a solitary flower (rarely in pairs), axillary; fruits strongly angled, triangular in cross section, tapering above, 3-locular 6. Merope
b. Inflorescence a raceme or panicle, few-to many-flowered, axillary or pseudoterminal; fruits globose, obtuse to rounded above, 5- or 4-locular 8. Pamburus
10a. Stamen. 4 or more times as many as the petals, often it regularly polyadelphous; ovary 8 - 20-locular with 4 or more ovules in each locule; fruit a hesperidium 3. Citrus
b. Stamens twice as many as the petals, often free (rarely polyadelphous); ovary 2 - 5 -locular with 1 or 2 ovules in each locule; fruit a juicy berry resembling small-sized lemons or oranges 11
11a. Leaves unifoliolate; petioles wingless; leaflet blades with many, close, parallel secondary veins connected with fine reticulate tertiary veinlets 2. Atalantia
b. Leaves tri- or unifoliolate; petioles winged; leaflet blades with fewer, prominulent secondary veins with inconspicuous veinlets 10. Pleiospermium

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