Herbs, thistle-like. Leaves alternate, toothed, lobed or pinnatisect, spinescent. Heads
homogamous, globose, ovoid or oblong, solitary or crowded, rarely corymbose. Flowers
white or purple, all similar, usually bisexual, tubular; limb cylindric, slender, usually
unequal, 5-fid. Involucral bracts many-seriate, appressed with long, erect, spreading or
recurved, spmous tips; Inner erect, slender, scanous. Receptacle flat, densely bristly.
Filaments glabrous; anther bases Sagittate; auricles connate; tails branched or villous.
Style arms narrow, short or long. Achenes, usually obovate or oblong, compressed,
glabrous, 5-many ribbed. Pappus hairs 1-seriate, slender or flattened, simple, free,
caducous, rarely absent.
E. Mediterranean to C. Asia; ca 400 species, 5 in India.
Literature.
RECHINGER, K.H. (1912) Flora Iranica. 90 (1,5): 1-329. SCHTEPA, 1.5.
(1962).
Palynological data for the Systematic; of the genus Cousinia Cass. Pollen et Spores 4(2):375.